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transverse zones There are three longitudinal zones the vermian, paravermian or intermediate, and lateral and there seems to be conSensory radiatons siderable overlapping from one to to areas 4 and 6 another Chambers and Sprague, on the basis of their investigations in Lateral ventral nucleus cats, concluded that the vermian zone coordinates movements of the eyes and body with respect to gravity and movement of the head in space The intermediate zone, which receives both peripheral and central projections (from motor cortex), in uences postural tone Rubrothalamic tract Dentatothalamic tract and also individual movements of the ipsilateral limbs The lateral zone is concerned mainly with coordination of movements of the ipMidbrain silateral limbs but is involved in Deep tegmental gray other functions as well The efferent bers of the cerParvocellular part Red ebellar cortex, which consist essennucleus Magnocellular part tially of the axons of Purkinje cells, project onto the deep cerebellar nuclei (see below) The projections Ventral tegmental decussation from Purkinje cells are inhibitory; those from the nuclei are excitDecussation of superior cerebellar peduncle Rubrobulbar tract atory According to the scheme of Rubrospinal tract Jansen and Brodal, cells of the verCerebellotegmental and cerebellorubral tract Tegmentospinal tract mis project mainly to the fastigial Superior cerebellar peduncle nucleus; those of the intermediate zone, to the globose and emboliform nuclei (represented by the interpositus nucleus in animals); and Emboliform Cerebellar nuclei those of the lateral zone, to the denDentate tate nucleus The deep cerebellar nuclei, in turn, project to the cerebral cortex and certain brainstem nuclei via two main pathways: bers from the dentate, emboliform, and globose nuclei form the superior cerebellar peduncle, enter the upper pontine tegmentum as the brachium conjunctivum, decussate the level of the inferior colliculus, Figure 5-2 Cerebellar projections to the red nucleus, thalamus, and cerebral cortex (Adapted by permis- and ascend to the ventrolateral nusion from House EL et al: A Systematic Approach to Neuroscience, 3rd ed New York, McGraw-Hill, cleus of the thalamus and, to a 1979) lesser extent, to the intralaminar thalamic nuclei (Fig 5-2) Some of the ascending bers, soon after their decussation, synapse in the pressure on the foot pad Ablation of a cerebellar hemisphere in red nucleus, but most of them traverse this nucleus without tercats and dogs yields inconsistent results, but in monkeys it causes minating Ventral thalamic nuclear groups that receive these ashypotonia and clumsiness of the ipsilateral limbs; if the dentate cending efferent bers project to the supplementary motor cortex nucleus is included in the hemispheric ablation, these abnormalities of that side Since the pathway from the cerebellar nuclei to the are more enduring and the limbs also show an ataxic or intention thalamus and then on to the motor cortex is crossed and the contremor nection from the motor cortex through the corticospinal is again As one might expect, cerebellar function and structure are crossed, the effects of a lesion in one cerebellar hemisphere are hardly as simple or precise as the preceding outline suggests The manifest by signs on the ipsilateral side of the body seminal studies of Chambers and Sprague and of Jansen and Brodal One pathway forms a loop, called the Mollaret triangle, that indicate that in respect to both its afferent and efferent projections, is of clinical interest (see also page 85) A small group of bers the cerebellum is organized into longitudinal (sagittal) rather than.

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